Santiria Blume

Dioecious trees, sometimes with plank buttresses or stilt roots; bark usually densely lenticellate; resin white to translucent. Leaves evergreen, exstipulate, imparipinnate or rarely unifoliolate, pulvinulus evident on terminal petiolule and usually on laterals; leaflets entire; intercostal tertiary veins alternate-percurrent and admedially ramified.  Inflorescences axillary or pseudoterminal, paniculate; male inflorescences generally longer than female. Receptacle shallowly concave; perianth 3-parted; calyx divided to varying depths; petals distinct, valvate at apex but imbricate along sides; stamens in 2 subequal or unequal series (sometimes the antepetalous series obsolete or vestigial), inserted on disk or at its base or rarely on the rim; anthers continuous with filaments or sagittate; pistillode reduced but containing reduced locules; disk intrastaminal, annular; gynoecium 2–3-locular, style short and thick, stigma (sub)sessile, 3-lobed or truncate.  Fruit a compound, indehiscent drupe, small (less than 2.5 cm long), oblique and ovoid or ellipsoid, rarely globose or acuminate; stigma markedly excentric, offset often more than 90º from vertical; calyx usually persistent; exocarp thin and almost smooth when dry; mesocarp thin; endocarp thinly cartilaginous; fruit 1—3-seeded, aborted locules strongly compressed, forming small, articulated, detachable plates on periphery of pyrene between the fruit base and excentric apex. Cotyledons 5-lobed or -fid, folded or contortuplicate.  Germination epigeal or hypogeal; first eophylls opposite and simple or pinnate.

About 22 spp., in W Africa, Malay Peninsula, Borneo, Sumatra, Philippines, Moluccas, Celebes, New Guinea.

Original Publication: Santiria Blume, Mus. Bot. Ludg. Bat. 1: 209, f. 40 (1850)

Synonyms: Trigonochlamys Hook.f. (1860).
Icicaster Ridl. (1917).

Bibliography: Leenhouts, Fl. Males. I, 5: 229–238 (1956).

Web Links:
Tropicos – Santiria
The Plant List – Santiria